Jerome Goddard

Jerome Goddard

Extension Professor

Medical / Veterinary Entomology

Mailing Address:
Box 9775
Mississippi State, MS 39762
  • Ph.D., Mississippi State University
  • M.S., University of Mississippi
  • B.A.E., University of Mississippi

Research Areas:
  • Ecology and epidemiology of tick-borne diseases

Extension Areas:
  • Public health entomology
  • Medical entomology
  • Management and control of insects and other arthropods which are harmful to humans and animals
  • Vector-borne diseases such as Lyme disease, Rocky Mountain spotted fever, and West Nile virus

Courses Taught:
  • EPP 4173/6173 - Medical and Veterinary Entomology
  • EPP 4313/6313 - Forensic Entomology


  • Goddard, J., G. De Jong, F. S. Meyer. 2020. Unidirectional en masse larval dispersal of blow flies. Food Webs 23:e00137. Download
  • Kooienga, E., J. Goddard. 2020. Maggot debridement therapy: a suitable treatment option for chronic wound care. Journal of the Mississippi State Medical Association 61:10-12.
  • Meyer, F. S., M. Monroe, H. Williams, J. Goddard. 2020. Solenopsis invicta x richteri necrophagous behavior causes post-mortem lesions in pigs which serve as oviposition sites for Diptera. Forensic Science International Reports 2:100067.  Download
  • Goddard, J., B. Layton. 2019. Spiders: Brown Recluse and Black Widow. Mississippi State University Extension Publication P2154. Download
  • Goddard, J., M. Allerdice, S. Portugal, G. Moraru, C. Paddock, J. G. King. 2019. What is going on with the genus Dermacentor? Hybridizations, introgressions, oh my! Journal of Medical Entomology. Download
  • Goddard, J., P. Stewart, 2019. Insect and other arthropod bites. Wolters Kluwer Publishers, Alphen aan den Rijn, South Holland. Download
  • Goddard, J., P. Stewart, H. Deerman, T. Nations, W. Varnado. 2018. Development and resolution of cutaneous lesions caused by black fly bites (Diptera: Simuliidae). American Journal of Medicine 131(10):e415-e416. Download
  • Hecht, J., M. Allerdice, E. Dykstra, L. Mastel, R. Eisen, T. Johnson, H. Gaff, A. Varela-Stokes, J. Goddard, B. Pagac, C. Paddock, S. Karpathy. 2019. Multistate survey of American dog ticks for Rickettsia species. Vector-Borne and Zoonotic Diseases 19(9):652-657.  Download
  • Lee, J., K. Grunnet, R. Flowers, B. Zlotoff, M. Noland, J. Goddard. 2019. Larval tick infestation causing an eruption of pruritic papules and pustules. Cutis 104:E1-E3. Download
  • McInnis, S., J. Goddard, H. Deerman, T. Nations, W. Varnado. 2019. Insecticide resistance testing of Culex quinquefasciatus and Aedes albopictus from Mississippi. Journal of the American Mosquito Control Association 35:147-150.
  • Portugal, S., M. Allerdice, G. Moraru, J. King, C. Paddock, T. Becker, T. Smith, J. Goddard. 2019. Molecular phylogeny of Dermacentor parumapertus from two locations within its geographical range. Journal of Medical Entomology 56(4):979-983.  Download
  • Campbell, Y., L., X. Zhang, W. Shao, J. B. Williams, T. J. Kim, J. Goddard, S. Abbar, T. W. Phillips, M. W. Schilling. 2018. Use of nets treated with food-grade coatings on dry-cured ham to control Tyrophagus putrescentiae infestations without impacting sensory properties. Journal of Stored Products Research 76:30-36. Download
  • Goddard, J., L. Lampton, 2018. Tick-borne diseases in Mississippi. Journal of the Mississippi State Medical Association 59:432-441.
  • Moraru, G., S. Portugal, A. Bednarova, S. McInnis, C. Paddock, T. Becker, T. Smith, Z. Svobodova, J. Goddard. 2018. Morphometric analysis of adult Dermacentor parumapertus from various locations within its geographical range. Journal of Medical Entomology 55(4):871-876.  Download
  • Nations, T. M., W. Varnado, A. Harrison-Lewis, H. Deerman, S. McInnis, J. Goddard. 2018. An annotated list of black flies occurring in Mississippi. Transactions of the American Entomological Society 144:539-550.  Download
  • Varnado, W., J. Goddard, 2018. A survey of environmental and behavioral aspects of human West Nile cases in Mississippi. Journal of the Mississippi State Medical Association 59:134-138.
  • Goddard, J. 2017. A crack-and-crevice application assay for selected insecticides to control the ham mite, Tyrophagus putrescentiae. Journal of Entomological Science 52(3):304-307. Download
  • Goddard, J. 2017. Not all erythema migrans lesions are Lyme disease. American Journal of Medicine 130(2):231-233. Download
  • Goddard, J., G. Moraru, S. McInnis, S. Portugal, D. Yee, H. Deerman, W. Varnado. 2017. A statewide survey for container-breeding mosquitoes in Mississippi. Journal of the American Mosquito Control Association 33:229-232. Download
  • Goddard, J., W. Varnado, 2017. Palpable purpura from chigger bites. Midsouth Entomologist 9: 77-78. Download
  • Outlaw, D., D. Larson, J. Goddard. 2017. Mosquito vectors of avian malaria in Mississippi: a first look. Journal of Parasitology. Download
  • Paddock, C., M. Allerdice, W. Nicholson, M. Levin, T. Smith, T. Becker, R. Delph, R. Knight, J. Ritter, J. Sanders, J. Goddard, 2017. Isolation and characterization of a unique strain of Rickettsia parkeri associated with the hard tick Dermacentor parumap ertus in the western United States. Applied and Environmental Microbiology. Download
  • Varnado, W., J. Goddard. 2017. A survey of environmental and behavioral aspects of human West Nile cases in Mississippi. Journal of the Mississippi State Medical Association 58:158-161.
  • Zhang, X., Y. Campbell, T. Phillips, S. Abbar, J. Goddard, M. W. Schilling, 2017. Application of food-grade ingredients to nets for dry cured hams to control mite infestation. Meat and Muscle Biology 1(1):53-60. Download
  • Goddard, J. 2016. Essential facts about mosquito control and Zika virus. American Journal of Medicine 129(11):1149-1152. Download
  • Goddard, J. 2016. Top 10 facts you should know about mosquito control and Zika virus. Journal of the Mississippi State Medical Association 57:178-180.
  • Goddard, J., M. Allerdice, C. Paddock, J. S. Portugal. 2016. Spotted fever group rickettsiae in immature Amblyomma maculatum from Mississippi (Acari: Ixodidae). Journal of Entomological Science 51(4):329-331. Download
  • Goddard, J., S. Portugal, K. Mascheck, 2016. Antibody titers to bed bug salivary proteins in chickens from an infested broiler breeder house. Midsouth Entomologist 9:1-4. Download
  • Goddard, J., W. C. Varnado, S. Hand, F. Meyer. 2016. Chikungunya in Mississippi: The Health Department Response to Imported Cases. Journal of the Mississippi State Medical Association 57(5): 1381141.
  • Goddard, J., W. C. Varnado, S. Hand, F. Meyer. 2016. Chikungunya in Mississippi: The Health Department Response to Imported Cases. Journal of the Mississippi State Medical Association 57: 138-140.
  • Nations, T., K. Edwards, J. Goddard. 2016. The George H. Bradley black fly papers (Diptera: Simuliidae). Midsouth Entomologist 8:73-75. Download
  • Portugal III, J. S., J. Goddard. 2016. Field observations of questing and dispersal by colonized nymphal Amblyomma maculatum. Journal of Parasitology 102(4):481-483. Download
  • Portugal, S., J. Goddard. 2016. Seasonality and distribution of immature Amblyomma maculatum (Acari: Ixodidae) in Mississippi: A review and new records. Journal of Entomological Science 51(1):1-8. Download
  • Portugal, S., J. Goddard. 2016. Survival of unfed Amblyomma americanum (Acari: Ixodidae) nymphs under laboratory conditions for two years. Journal of Entomological Science 51: 98-99. Download
  • Varnado, W., J. Goddard. 2016. Use of the VectorTest for advanced warning of human West Nile viruscases in Mississippi. Journal of Environmental Health 79: 20-24. Download
  • Yoder, J. A., B. W. Nelson, C. R. Randazzo, J. S. Portugal III, J. Goddard. 2016. Dehydration tolerance in relation to short day length in nonfed Amblyomma maculatum Koch, 1844 ticks (Acari: Ixodidae). International Journal of Acarology 43(2):173-179. Download
  • Goddard, J., A. Moraru, 2015. Philosophical considerations of quantum mechanics. Journal of the Mississippi Academy of Sciences 60:336-338.
  • Goddard, J., C. Ward, J. Dagg, M. Mendonca, R. G. Robbins. 2015. New records for Amblyomma rotundatum Koch from the cane toad, Rhinella marina L. in Florida, with notes on identification of the nymphs. Systematic and Applied Acarology 20(5):456-467. Download
  • Goddard, J., J. S. Portugal III. 2015. Cutaneous lesions due to bites by larval Amblyomma americanum ticks. JAMA Dermatology 151(12):1373-1375. Download
  • Goddard, J., J. Seltzer, F. S. Meyer. 2015. New state record for Chrysomya ruffifacies in Mississippi. Journal of Entomological Science 50:67-68. Download
  • Goddard, J., K. Maschak. 2015. Laboratory assays with various insecticides against bed bugs taken from a poultry house in Mississippi. Midsouth Entomologist 8:10-15. Download
  • Goddard, J., M. A. Caprio, J. Goddard II. 2015. Diffusion rates and dispersal patterns of unfed versus recently fed bed bugs (Cimex lectularius L.). Insects 6(4):792-804. Download
  • Goddard, J., M. Embers, A. Hojgaard, J. Piesman. 2015. Comparison of tick feeding success and vector competence for Borrelia burgdorferi among immature Ixodes scapularis of both southern and northern clades. Journal of Medical Entomology 52(1):81-85. Download
  • Paddock, C. D., J. Goddard. 2015. The evolving medical and veterinary importance of the Gulf Coast tick. Journal of Medical Entomology 52(1):230-252. Download
  • Portugal III, J. S., J. Goddard. 2015. Attempts to feed larval Amblyomma americanum (L.) (Acari: Ixodidae) on three different arthropod hosts. Journal of Vector Ecology 40(1):202-204. Download
  • Portugal III, J. S., J. Goddard. 2015. Collections of immature Amblyomma maculatum from Mississippi. Systematic and Applied Acarology 20(1):20-24. Download
  • Portugal III, J. S., J. Goddard. 2015. Evaluation of human attachment by larval Amblyomma maculatum. Journal of Medical Entomology 53(2):451-453. Download
  • Varnado, W. C., J. Goddard. 2015. Abundance and diversity of mosquito species collected from a rural area of central Mississippi: implications for West Nile virus transmission in Mississippi. Journal of the American Mosquito Control Association 31:182-186. Download
  • Ekenna, O., C. D. Paddock, J. Goddard. 2014. Gulf coast tick rash illness in Mississippi caused by Rickettsia parkeri. Journal of the Mississippi State Medical Association 55(7):216-219.
  • Ferrari, F. A. G., J. Goddard, C. D. Paddock, A. Varela-Stokes. 2014. Ultrastructure of presumed "Candidatus Rickettsia andeanae" in Amblyomma maculatum. Journal of Medical Entomology 51(6): 1317-1321. Download
  • Goddard, J. 2014. Long-term efficacy of various natural or "green" insecticides against bed bugs: a double-blind study. Insects 5(4):942-951. Download
  • Goddard, J., C. D. Paddock, 2014. Variability in ornamentation of adult Dermacentor parumapertus: implications for tick identification. Systematic and Applied Acarology 19(2):253-256. Download
  • Goddard, J., K. Hoppens, K. Lynn. 2014. Case report on enteric myiasis in Mississippi. Journal of the Mississippi State Medical Association 55(4):132-133.
  • Goddard, J., L. E. Goltz, K. T. Edwards, W. Smith, A. Varela-Stokes. 2014. Survey of adult Dermacentor variabilis collected in northern Mississippi for spotted fever group rickettsiae. Midsouth Entomologist 7:8-14. Download
  • Jones, K. H., N. Johnson, S. Yang, J. Stokes, W. Smith, R. Wills, J. Goddard, A. Varela-Stokes. 2014. Investigations into outbreaks of black fly attacks and subsequent avian haemosporidians in backyard-type poultry and other exposed avian species. Avian Diseases 59:24-30. Download
  • Sakamoto, J., J. Goddard, J. Ragson. 2014. Population and demographic structure of Ixodes scapularis in the eastern United States. PLoS One 9(7):e101389. Download
  • Ferrari, F.A., J. Goddard, G. Moraru, W.E. Smith, A.S. Varela-Stokes. 2013. Isolation of "Candidatus Rickettsia andeanae" in embryonic cells of naturally infected Amblyomma maculatum. Journal of Medical Entomology, 50(5):1118-1125. Download
  • Ferrari, F.A., J. Goddard, M.A. Caprio, C.D. Paddock, T. Mixson-Hayden, A.S. Varela-Stokes. 2013. Population analyses of Amblyomma maculatum ticks and Rickettsia parkeri using singlestrand confirmation polymorphism. Ticks and Tick-borne Diseases. Download
  • Goddard, J. 2013. Laboratory assays of various insecticides against bed bugs (hemiptera: Cimicidae) and their eggs. Journal of Entomological Science, 48(1):65-69. Download
  • Goddard, J., K.T. Edwards. 2013. Effects of bed bug saliva on human skin. Journal of the American Medical Association, 149(3):372-373. Download
  • Goddard, J., L. Goltz, R.G. Robbins. 2013. New records of immature Ixodes brunneus Koch from Mississippi. Journal of Vector Ecology, 38(2):415-416. Download
  • Goddard, J., N. Hasenkampf, K.T. Edwards, R.D. de Shazo, M. Embers. 2013. Bed bug saliva causes release of monocytic inflammatory mediators: plausible cause of cutaneous bite reactions. International Archives of Allergy and Immunology, 161:127-130. Download
  • Goltz, L., A.S. Varela-Stokes, J. Goddard. 2013. Survey of adult Ixodes scapularis say for disease agents in Mississippi. Journal of Vector Ecology, 38(2):401-403. Download
  • Goltz, L., J. Goddard. 2013. Observations on the seasonality of Ixodes scapularis say in Mississippi. Systematic & Applied Acarology, 18:212-217. Download
  • Moraru, G., J. Goddard, A. Murphy, D. Link, J. Belant, A.S. Varela-Stokes. 2013. Evidence of antibodies to spotted fever group rickettsiae in small mammals and quail from Mississippi. Vector-borne and Zoonotic Disease, 13:1-5. Download
  • Moraru, G., J. Goddard, C.D. Paddock, A.S. Varela-Stokes. 2013. Experimental infection of cotton rats and bob white quail with Rickettsia parkeri. Parasites and Vectors, 6:70. Download
  • Plotkin, D., J. Goddard. 2013. Blood, sweat, and tears: a review of the Hematophagous, Sudophagous, and Lachryphagous Lepidoptera. Journal of Vector Ecology, 38(2):289-294. Download
  • de Shazo, R.D., M.F. Feldlaufer, M.C. Mihm, J. Goddard. 2012. Bullous reactions to bed bug bites reflect cutaneous vasculitis. American Journal of Medicine 125(7):688-694. Download
  • Ferrari, F.A., J. Goddard, C.D. Paddock, A.S. Varela-Stokes. 2012. Rickettsia parkeri and Candidatus Rickettsia andeanae in Gulf Coast ticks, Mississippi, USA. Emerging Infectious Diseases, Vol. 18:1705-1707. Download
  • Goddard, J. 2012. Bed bug bites: everything you need to know but were afraid to ask. UBM Medica Psychiatric Times. Download
  • Goddard, J. 2012. Physician's Guide to Arthropods of Medical Importance. Sixth Edition. Taylor and Francis Publishers (CRC Press), 515 pp. Download
  • Goddard, J. 2012. Public Health Entomology. Taylor and Francis Publishers (CRC Press), 282 pp.
  • Goddard, J., A.S. Varela-Stokes, R. Finley. 2012. Lyme Disease-like Illness in the South. Journal of the Mississippi State Medical Association 53(3):68-72. Download
  • Goddard, J., A.S. Varela-Stokes, W.E. Smith, K.T. Edwards. 2012. Artificial infection of the bed bug with Rickettsia parkeri. Journal of Medical Entomology 49(4):922-926. Download
  • Goddard, J., D.E. Fleming, J.L. Seltzer, S.H. Anderson, C. Chesnut, M. Cook, E.L. Davis, B. Lyle, S. Miller, E.A. Sansevere, W. Schubert. 2012. Insect succession on pig carrion in north-central Mississippi. Midsouth Entomologist 5:39-53. Download
  • Goddard, J., G.T. Baker, F.A. Ferrari, C. Ferrari. 2012. Bed bugs and bat bugs: a confusing issue. Outlooks on Pest Management 23:125-127. Download
  • Goddard, J., L. Bircham, and R.G. Robbins. 2012. New tick records from a mink, Neovison vison (Schreber), in Mississippi. Systematic & Applied Acarology 17(3):251-253. Download
  • Goddard, J., R. deShazo. 2012. Psychological effects of bed bug attacks (Cimex lectularius). American Journal of Medicine 125(1):101-103. Download
  • Hill, J.G., J. Goddard. 2012. Medical and veterinary importance of grasshoppers, katydids, and crickets (Insecta: Orthoptera). Journal of the Mississippi Academy of Sciences 57:172-177. Download
  • Moraru, G., J. Goddard, A.S. Varela-Stokes. 2012. Observations on host preference and feeding success of immature amblyomma maculatum. Journal of Entomological Science 47:221-226. Download
  • Thorn, M., W. Varnado, J. Goddard. 2012. First record of Aedes japonicas in Mississippi. Journal of the American Mosquito Control Association 28(1):43-44. Download
  • Edwards, K.T., A.S. Varela-Stokes, C.D. Paddock, J. Goddard. 2011. Gotch ear in a goat. Journal of Vector-Borne and Zoonotic Diseases 11:1217-1219. Download
  • Edwards, K.T., A.S. Varela-Stokes, J. Goddard. 2011. Distribution of spotted fever group Rickettsiae in select tissues of experimentally-infected and field-collected gulf coast ticks. Journal of Medical Entomology 48(3):687-690. Download
  • Edwards, K.T., J. Goddard, T.L. Jones, C.D. Paddock, A.S. Varela-Stokes. 2011. Cattle and the Natural History of Rickettsia parkeri in Mississippi. Journal of Vector-Borne and Zoonotic Diseases 11(5):485-491. Download
  • Goddard, J. 2011. Arthropods, tongue worms, leeches, and mihropod-borne diseases. IN: Tropical Infectious Diseases, 3rd Edition, Guerrant, R.L., Walker, D.H., and Weller, P.F., Eds., Elsevier Publishers, New York, pp. 868-878. Download
  • Goddard, J. 2011. Bed bugs (Cimex lectularius L.) in Mississippi: survey of the scope, extent, and control of the problem. Midsouth Entomologist 4(4):57-62. Download
  • Goddard, J. 2011. Ectoparasites, part two: mites and ticks. IN: The Mallis Handbook of Pest Control, 10th Edition, GIE Media, Cleveland, OH, pp. 551-585.
  • Goddard, J., A.S. Varela-Stokes, J.C. Schneider. 2011. Observations on questing activity of Adult Gulf Coast Tick, Amblyomma maculatum Koch (Acari: Ixodidae), in Mississippi, U.S.A. Systematic & Applied Acarology 16:195-200. Download
  • Goddard, J., J. Arrigoni. 2011. Temporary vision improvement and pain relief after insect sting. Journal of Emergency Medicine. Download
  • Goddard, J., K. Hataway. 2011. Malaria in Mississippi: History, epidemiology, and current status. Journal of the Mississippi Academy of Sciences 56(4):223-332. Download
  • Goddard, J., K.T. Edwards, R. deShazo. 2011. Observations on development of cutaneous lesions from bites by the common bed bug, Cimex lectularius L. Midsouth Entomologist 4(2):49-52. Download
  • Goddard, J., L. Bircham, R.G. Robbins. 2011. New tick-host records from Mississippi, U.S.A. Systematic & Applied Acarology 16:212-214. Download
  • Edwards, K.T., J. Goddard, A.S. Varela-Stokes, M.B. Layton, C.L. Huston, L.D. Foil. 2010. Current topics in medical and veterinary entomology: results of a roundtable discussion. Midsouth Entomologist 3(1):67-72. Download
  • Goddard, J., A.S. Varela-Stokes. 2009. The discovery and pursuit of American Boutonneus Fever: a new spotted fever group rickettsia. Midsouth Entomologist, Vol. 2(1):47-52. Download
  • Goddard, J., J. Goddard II, X. Wang. 2009. Proportion of adult lone star ticks (Amblyomma americanum) questing in a tick population. Journal of Mississippi Academy of Sciences 54:206-209. Download
  • Goddard, J., R. deShazo. 2009. Multiple feeding by the common bed bugs, Cimex lectularius, without sensitization. Midsouth Entomologist 2(2):90-92. Download